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This work has been done within the efforts of the European Reference Network EpiCARE ( Conflict of interest The authors declare that the research was conducted in the absence of any commercial or financial relationships that could be construed as a potential conflict of interest

Quiescin-sulfhydryl oxidase 1 (QSOX1) is a sulfhydryl oxidase located in the GA that catalyzes the oxidation of thiol groups to form disulfide bonds in proteins, accompanied by the reduction of O 2 to H 2 O 2 [116, 117]