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In the triggering step of RQC, Hel2 forms a K63-linked polyubiquitin chain at lysine residues K6 or K8 of ribosome protein uS10 encoded by RPS20 around the colliding interface

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Subsequently, the Cys-Gly dipeptide is degraded by membrane-bound Anpep to yield cysteine and glycine for another glutathione cycle