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Lens cells need to continually extrude intracellular protons, which accumulate due to inward movement of positive ions from the extracellular space and lactic acid from anaerobic glycolysis

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Moreover, the stiffness of the extracellular matrix can influence DNA methylation through protein kinase C -dependent nuclear transport of DNMT3L, thereby connecting biomechanical signals to the epigenetic regulation of stem cell pluripotency