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Although mitochondria have antioxidant defenses like SOD, which helps convert superoxide anions into less harmful molecules, too much mitochondrial ROS can cause oxidative stress and harm both the mitochondria and the sperm cell

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Tudor (Tud), the familys founder member and one of the 23 Tudor domain-containing proteins discovered in Drosophila, is crucial for the correct localization of Aub in the germline and for the assembly of germplasm [420, 421]