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Regarding the mechanisms of crossing the nasopharyngeal barrier, a combination of transcellular and paracellular pathways seems possible if meningococci using the transcellular pathway induce a sufficient level of inflammation to alter epithelial permeability, allowing other meningococci to follow the paracellular pathway

doi:10.1007/s11064-012-0895-x Lin JB, Apte RS

nucleotide binding then brings the NBDs closer together, reducing the inter-NBD distance to ~30