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The math, the chemistry, and the research framing are all below

DNA methylation in promoter regions can suppress or activate carotenoid-related genes (e.g., hypomethylation of the PSY1 promoter is associated with increased lycopene accumulation in ripening tomatoes) (2015a)

However, the activity reported was very low, which could be enhanced to 963-fold for a triple mutant E423Y/E442Q/D445N of BsGGT, developed by site-directed and random mutagenesis approach (Suzuki et al., 2010)

Taken together, these studies paint a complex picture of the role of Skp2 in cell cycle progression but suggest an important role in proliferation and likely in the pathogenesis of cancer

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